As the Covers start to kidnap citizens, including Mako, using their life force to power themselves, Satsuki triggers an explosion to allow Ryuko to escape. One month later, the Elite Four have joined up with Nudist Beach to fight against the Covers as they take over Japan, while Ryuko has remained in a coma since the battle. Meanwhile, Satsuki has been held captive by Ragyo, who explains that her father, Soichiro, had faked his own death and disguised himself as Isshin in order to raise Ryuko and prepare his revenge against her.